Schematic drawing showing the general organization and development of E. multilocularis metacestodes. (A) Early brood capsule bud. (B) Brood capsule with protoscolex bud. (C) Brood capsule with protoscolex in late development. (D) Brood capsule with invaginated protoscolex. The syncitial tegument is shown in orange, the germinative cells in brown, glycogen/lipid storage cells in violet, calcareous corpuscle cells in light blue, nerve cells in green and muscle cells and fibers in red. bc, brood capsule; GL, germinal layer; HF, hydatid fluid; LL, laminated layer; ps, protoscolex; r, rostellum; s, sucker.